Donald A. McFarlane
Professor of Environmental Science
Professor of Biology
Office: Keck Science Center 43
Phone: 909-607-2564
Office Hours:
Web Site:
Educational Background:
BSC. (Honours) Zoology; University of Liverpool, 1977. MSc. Zoology. Queen's University of Belfast, 1982. PhD. Ecology. University of Southern California, 1987.
Courses Taught:
Ecology and Evolution.
Research Interests:
The ecology of extinction. Late Quaternary paleobiology and paleoecology. Ecology of cave ecosystems. Ecology of bats. Evolutionary biogeography of tropical islands. Mammalogy. Cave paleontology.
Thesis Topics:
**** See my faculty web-page for current opportunities *****
Studies in evolution of islands, cave biology
Late Quaternary paleontology of mammals; recent mammalian extinctions
Island biogeographic analyses
Late Quaternary climate and sea-level change.
Cave science, including microclimate modelling.
Selected Publications List: Click to open new window.
1.   Lundberg, J. and McFarlane, D. A. . (2008). Kents Cavern. A field guide to the natural history..  William Pengelly Cave Studies Trust. Buckfastleigh. : 44p..
2.   Wurster C. M., Patterson W.P., McFarlane D.A., Wassenaar L. I., Hobson K.A., Athfield N. B., and Bird M. I. . (2008). Stable carbon and hydrogen isotopes from bat guano in the Grand Canyon, USA, reveal Younger Dryas and 8.2 ka events.. Geology 36:683-686.   36: 683-686.
3.   Christenson K. and McFarlane D. A. . (2007). An ecologically significant range extension for Hahn’s Short-tailed fruit bat, (Carollia subrufa) from southwestern Costa . Chiroptera Neotropical   13(2): 319-321.
4.   Wurster, C. M., McFarlane, D. A., and Bird, M. I. . (2007). Spatial and temporal expression of vegetation and atmospheric variability from stable carbon and nitrogen isotope analysis of bat guano in the southern United States.. Geochimica et Cosmochimica Acta   71: 3302-3310. Abstract
5.   Lundberg, J. and McFarlane, D.A. . (2007). Speleogenesis of the Mt. Elgon ‘Elephant’ Caves, Kenya.. Geological Society of America Special Paper 404 Harmon, R.S. and Wicks, C.  : 51-63. Abstract
6.   Lundberg, J. and McFarlane, D.A. . (2007). Pleistocene depositional history in a periglacial terrane: A 500ka record from Kents Cavern, Devon, United Kingdom.. Geosphere   3: 199-219. Abstract
7.   Lundberg, J. and McFarlane, D.A. . (2006). A minimum age for canyon incision and for the extinct Molossid bat, Tadarida constantinei, from Carlsbad Caverns National Park, New Mexico.. Journal of Cave and Karst Studies   68(3): 155-177.
8.   McFarlane D. A. and Blake, J. . (2005). The Late Pleistocene Hutias (Geocapromys brownii) of Red Hills Fissure, Jamaica.. Geological Journal   40: 399-404. Abstract
9.   McFarlane D. A. and Lundberg, J. . (2005). The 19th Century excavation of Kent’s Cavern, England.. Journal of Cave and Karst Studies   67(1): 39-47. Abstract
10.   McFarlane, D. A. . (2004). Guano. Encyclopedia of Caves and Karst Science (J. Gunn, ed.) Fitzroy Dearborn, New York : 409-410.
11.   McFarlane, D. A., Lundberg, J. . (2004). An unusual lava cave from Ol Doinyo lengai, Tanzania.. Journal of Cave and Karst Studies   66(3): 98-101. Abstract
12.   McFarlane, D. A. and Lundberg . (2004). Reliquiae Diluviane Alter: Last Interglacial flood deposits in caves of the West Indies. Studies of Cave Sediments  Kluwer/Plenum Press, New York : 313-322.
13.   McFarlane D. A., Lundberg J. . (2002). A middle Pleistocene age and biogeography for the extinct rodent Megalomys curazensis from Curacao, Netherlands Antilles. Caribbean Journal of Science   38: 278-281. Article
14.   Rega, E., McFarlane, D. A., Lundberg, J. and Christenson, K . (2002). A new Megalonychid sloth from the Late Wisconsinan of the Dominican Republic.. Caribbean Journal of Science   38: 11-19. Article
15.   McFarlane D. A., Lundberg J., and Fincham A. G . (2002). A late Quaternary paleoecological record from caves of southern Jamaica, West Indies. Journal of Cave and Karst Studies   64(2): 117-125. Abstract Article
16.   McFarlane D. A. and Debrot, A. . (2001). An extinct Oryzomyine rodent from the Quaternary of Curaçao, Netherlands Antilles. Caribbean Journal of Science   37: 182-184. Article
17.   Nieves-Rivera, A. and McFarlane D. A. . (2001). In search of the extinct hutia in cave deposits of Isla de Mona. National Speleological Society News   59(4): 92-95.
18.   Lugo A. E. et al . (2001). Puerto Rican Karst – A Vital Resource. United States Dept. of Agriculture. Forest Service. Technical Report   WO-65: 107. Article
19.   McFarlane D.A., Vale A., Christenson K., Lundberg J., Atilles G., and Lauritzen S.E. . (2000). New specimens of Late Quaternary extinct mammals from caves in Sanchez Ramirez Province, Dominican Republic. Caribbean Journal of Science   36: 163-166. Article
20.   McFarlane, D. A., J. Lundberg, and D. C. Ford . (2000). The age of the Woolly Rhino from Dream Cave, Derbyshire, UK. Cave and Karst Science   37(1): 25-28.
21.   McFarlane, D.A. . (1999). A note on sexual dimorphism in Nesophontes edithae (Mammalia: Insectivora), an extinct island-shrew from Puerto Rico. Caribbean Journal of Science   35: 142-143. Article
22.   McFarlane D. A. . (1999). Late Quaternary Fossil Mammals and Last Occurrence dates from Caves at Barahona, Puerto Rico. Caribbean Journal of Science   35: 238-248. Article
23.   McFarlane D.A. . (1999). A comparison of methods for the probabilistic determination of vertebrate extinction chronologies. Extinctions in Near Time: Causes, Contexts, and Consequences (R. D. E. MacPhee, ed.) Plenum Press, New York : 95-106.
24.   McFarlane D. A., Lundberg J., Flemming, C. E., MacPhee R.D.E., and Lauritzen S.E. . (1998). A second Pre-Wisconsinan locality for the extinct Jamaican rodent, Clidomys (Rodentia: Heptaxodontidae). Caribbean Journal of Science   34: 315-317. Article
25.   Flemming C. and McFarlane D. A. . (1998). New Caribbean locality for the extinct Great White Shark Carcharodon. Caribbean Journal of Science   34: 315-317. Article
26.   McFarlane, D. A. and Ford, D. C. . (1998). The age of the Kirkdale Cave palaeofauna. Cave and Karst Science   25(1): 3-6.
27.   Panuska, B. C., Mylroie, J. M., Armentrout, D. and McFarlane, D. A. . (1998). Magnetostratigraphy of Cueva Aleman, Isla de Mona, Puerto Rico and the species duration of Audubon’s Shearwater. Journal of Cave and Karst Studies   60(2): 96-100. Abstract Article
28.   Flemming, C. MacPhee, R. D. E., and McFarlane, D. A. . (1998). Thrice bitten: Late Quaternary mammal extinctions in the continental and insular New World. Journal of Vertebrate Paleontology   18(3;suppl):21A (abstract): .
29.   McFarlane D. A., MacPhee R.D.E., and Ford D. . (1998). Body size variability and a Sangamonian extinction model for Amblyrhiza, a West Indian megafaunal rodent. Quaternary Research   50: 80-89.
30.   McFarlane, D. A. . (1997). Jamaican cave vertebrates. Jamaica Underground. The caves, sinkholes and underground rivers of the island (Fincham, A. G., ed.) The Press, University of the West Indies : 57-62.
31.   McFarlane D.A., Keeler R.C., and Mizutani H. . (1995). Ammonia volatization in a Mexican bat cave ecosystem. Biogeochemistry   30: 1-8.
32.   Nieves-Rivera A.M., Mylroie J.E., and McFarlane D.A. . (1995). Bones of Puffinus Lherminieri Lesson (Aves: Procellaridae) and two other vertebrates from Cueva del Agua, Mona Island, Puerto Rico (West Indies) . Nat. Speleol. Soc. Bull.   57: 99-102.
33.   Rinehart R.B. and McFarlane D.A . (1995). An early Holocene vegetation record from the Salton Basin, California USA. Quaternary Research   43: 259-262.
34.   MacPhee R.D.E., McFarlane D.A., Arrendondo O., and Jiminez Vasquez O. . (1994). West Indian monkeys: new fossils and interpretations. American Journal of Physical Anthropology   Suppl. 18: 133. Article
35.   McFarlane D.A. and MacPhee R.D.E. . (1994). Amblyrhiza and the vertebrate paleontology of Anguillean caves. Boletin Soc. Venezolana Espel   27: 33-38.
36.   Biknevicius A., McFarlane D.A. and MacPhee R.D.E. . (1993). Body size in Amblyrhiza inundata (Rodentia; Caviomorpha) an extinct megafaunal rodent from the Anguilla Bank, West Indies: Estimates and implications. American Museum Novitates   3079: 1-25. Article
37.   Mizutani H., McFarlane D.A. and Kabaya Y. . (1992). Carbon and nitrogen isotopic signatures of bat guanos as a record of past environments. Mass Spectrometry   411: 67-82.
38.   Mizutani H., McFarlane D.A., and Kabaya Y. . (1992). Nitrogen and carbon isotope studies of a bat guano core from Eagle Creek Cave, Arizona, USA. Mass Spectrometry   40(1): 57-65.
39.   McFarlane D.A. . (1991). The species-genus relationship in Antillean bat communities. Mammalia   55(3): 363-370.
40.   McFarlane D.A. . (1991). The search for Anguilla's giant rodent. Terra   30(2): 34-39.
41.   MacPhee R.D.E., Lundberg J., McFarlane D.A., Biknevicius A., and Ford D.C. . (1990). New specimens of Amblyrhiza inundata and their bearing on its morphology, adaptations, and age. Proc. Congresso Intl. 50 Aniv Soc. Espeleolog Cuba   (Havana, January 1990): .
42.   McFarlane D.A. and Keeler R. . (1990). A proxy record of the Free-Tailed Bat population at Eagle Creek, Arizona. In. Proceedings of the Symposium on Managing Wildlife in the Southwest   Wildlife Society, Tucson, Arizona: 100-106.
43.   McFarlane D.A. . (199). Patterns of species co-occurrence in the Antillean bat fauna.. Mammalia   53: 59-66.
44.   McFarlane D.A. . (1989). Histoplasmosis and the proposed development of The Fountain, Anguilla, British West Indies. In. A Study of Fountain National Park and Fountain Cavern, Anguilla, British West Indies  National Speleological Foundation, Closter, NJ : 21-22.
45.   MacPhee R.D.E. and McFarlane D.A. . (1989). Amblyrhiza inundata, the giant "rat" of of Anguillae. Anguilla Life   2(3): 19-20.
46.   McFarlane D.A. and MacPhee R.D.E. . (1989). Amblyrhiza and the Quaternary bone caves of Anguilla, British West Indies. Cave Science   16: 31-34.
47.   McFarlane D.A. and Stager K.E. . (1989). An abbreviated catalogue of the Australian bats in the collections of the Natural History Museum of Los Angeles County, California, USA.. Macroderma   4: 72-76.
48.   MacPhee R.D.E., Ford D., and McFarlane D.A. . (1989). Pre-Wisconsinan mammals from Jamaica and models of late Quaternary extinction in the Greater Antillles. Quaternary Research   31: 94-106.
49.   Blood B.R. and McFarlane D.A. . (1988). A new method of calculating the wing area of bats. . Mammalia   52: 600-603.
50.   McFarlane D.A. . (1988). Endangered Cave Species. National Speleological Society News   46: 36-37.
51.   McFarlane D.A. . (1988). Amblyrhiza rediscovered on Anguilla. Newsletter, Paleontology Section of the National Speleological Society   3: 1-2.
52.   Blood B.R. and McFarlane D.A. . (1988). Notes on Vespertilionid bats from northern Thailand, with comments on the subgeneric status of Myotis altarum. Zeitschrift fur Saugetierkunde   53: 276-280.
53.   McFarlane D.A. . (1987). Radiant darkness - the many facets of the caves of Jackson's Bay, Jamaica.. Terra   25(6): 24-26.
54.   McFarlane D.A. . (1987). An annotated bibliography of Irish Bats. part 1. 1830-1900. The Leisler  Ulster Museum 3: 17-22.
55.   McFarlane D.A. . (1986). A late Quaternary mammal fauna and palaeoclimatic record from the Jackson's Bay caves, Jamaica. Annual Report of the Cave Research Foundation   1985: 21.
56.   McFarlane D.A. . (1986). Fungal isolates from the air of a Jamaican bat cave. North American Biospeleology Newsletter   33: 8.
57.   McFarlane D.A. . (1986). Cave bats in Jamaica. Oryx   20(1): 27-30.
58.   McFarlane D.A. and Blood B.R. . (1986). Taxonomic notes on a collection of Rhinolophidae Chiroptera) from northern Thailand, with a description of a new subspecies. Zeitschrift fur Saugetierkunde   51: 218-223.
59.   McFarlane D.A. . (1985). Transmission of Histoplasmosis in guano caves, North American. Biospeleology Newsletter   32: 7-8.
60.   McFarlane D.A. and Gledhill R.E. . (1985). The Quaternary bone caves and associated sites at Wallingford, Jamaica. Cave Science   12(3): 127-128.
61.   McFarlane D.A. . (1985). The rat-bat caves of Jamaica. Terra   23(3): 14-17.
62.   McFarlane D.A. . (1981). The design of oxygen rebreather equipment for use in foul-air speleology. Transactions of the British cave Research Association   8(3): 130-134.
63.   McFarlane D.A. and Garrett K.L. . (198). The prey of common barn owls (Tyto alba) in dry limestone scrub forest of southern Jamaica. Caribbean Journal of Science   25(1-2): 21-23. Article
64.   McFarlane D.A. . (1977). Some abandoned resurgence caves in the Betzula area. Cave Notes   21: 37-39.
65.   McFarlane D.A. . (1977). The Gouffre d'Arphidia. Cave Notes (Bristol Exploration Club Report)   21: 2-5.
66.   DeNault L.K. and McFarlane D.A. . (1195). Altruistic blood sharing between male vampire bats, Desmodus rotundus. Animal Behaviour   49: 855-856. Article
67.   McFarlane D.A. . (). Liverpool University expedition to Jamaica. Transactions of the British Cave Research Association   7(3): 150-168.